Alfred Knudson Jr., M.D.,Ph.D.

Fox Chase Cancer Center, Philadelphia, Pennsylvania, USA

"for his development of a two-hit mutational hypothesis for the origin of cancer that related hereditary and non-hereditary forms of retinoblastoma and other tumors; this hypothesis has become a paradigm for mutational theories on the origins of cancer and led directly to the concept of suppressor oncogenes."

Awards:

1997 Gairdner International Awardee