Oliver Smithies

† 1925-2017
Oliver Smithies received the Canada Gairdner International Award in 1990 for the discovery, development and application of gel electrophoresis methods that allow the separation and identification of specific proteins and nucleic acids.
Oliver Smithies also received the Canada Gairdner International Award in 1993 for pioneering work in the use of homologous recombination to generate targeted mutations in the mouse.